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, and this view agrees perfectly with all the details of distribution. The centre of distribution lies in Northern Asia, or in Arctic North America. From there the great genus _Lagopus_ spread east and west, reaching Europe by these vastly divergent routes at a time when the physical geography was very different from what it is to-day. Several of the species common to the Alps and Scandinavia have migrated from Siberia direct to Eastern Europe. But we can now imagine how from a similar centre in Asia--perhaps at a rather more remote time--a species spread eastward across North America and Greenland to Scandinavia, and westward along the mountain ranges of the Tian Shan and the mountains of Asia Minor to Greece, and finally to the Alps. We should then have the same species in the Alps and in Scandinavia, not far removed from one another; but how different were their paths of migration! This, however, is not an imaginary instance. Such a migration must have actually taken place in a good number of instances among the terrestrial invertebrates and also among plants. The view still current among many zoologists and botanists, that animals and plants were driven down into the plain from the mountains of Europe during the height of the Glacial period and there lived together till the return of a more genial temperature, when they retreated to their mountain homes, is a very plausible one. During their sojourn in the plain, the plants and animals--say from Scandinavia--intermingled with those from the Alps; and when the time of separation came, many Alpine forms retired northward with the Scandinavians, while many Scandinavians would go with the Alpines to their home. In this way the similarity between the Alpine and Scandinavian faunas and floras is assumed to have been brought about. These theories, first promulgated by Edward Forbes, were hailed with general satisfaction by the scientific world. Even Darwin says of them (p. 331), that grounded as they are on the perfectly well-ascertained occurrence of a former Glacial period, they seemed to him to explain in a satisfactory manner the present distribution of the Alpine and Arctic productions of Europe. To the present day this view meets with much favour among naturalists. It is somewhat similar to one which has recently been strongly supported by Professor Nehring and accepted by Professor Th. Studer and many others. They have never made it quite clear whether the pre-glaci
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