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of their origin in a dry and intensely cold climate? I quite agree with the views as to the Asiatic origin of the bulk of the Alpine flora, while the dry state of the Siberian climate is certainly indicated by the extremely feeble development of the glaciers during a large part of the Glacial period. We know, however, that in Pliocene and even in early Glacial times the atmospheric conditions must have been very different in Siberia. A great slice of Central Asia was under water, and numerous freshwater lakes covered the lowlands in the north, so that the climate must have been damp though not cold enough for the formation of extensive glaciers. Everything, in fact, seems to indicate that the migration of the Asiatic Alpine flora took place at a very early date--probably long before the Glacial period--either by the Oriental or by the Arctic route _via_ North America, Greenland, and Scandinavia. But would this not necessitate a survival of the Alpine plants in the Alps themselves? That is the view which has already been expressed with regard to the fauna, and the flora probably followed a very similar course. This is by no means a novel theory, however, and though unfortunately an untimely death has removed one of our very best authorities on the Alpine flora before he had completed his life's work, we have some indications in the earlier writings of John Ball that his opinions on the origin of that flora did not coincide with those held by the leading continental authors. To quote the words of this distinguished botanist (p. 576): "Is it credible that in the short interval since the close of the Glacial period hundreds of very distinct species and several genera have been developed in the Alps, and--what is no less hard to conceive--that several of these non-Arctic species and genera should still more recently have been distributed at wide intervals throughout a discontinuous chain some 1500 miles in length, from the Pyrenees to the Eastern Carpathians? Nor would the difficulties cease there. You would have left unexplained the fact that many of the non-Arctic types which are present in the Alps are represented in the mountains of distant regions, not by the same, but by allied species, which must have descended from a common ancestor; that one species of _Wulfenia_, for example, inhabits one small corner of the Alps, that another is found in Northern Syria, while a third allied species has its home in the Himalaya." Mr.
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