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ignificance to the differentiation of its alimentary canal into stomach and intestine. That in the higher _Annulosa_, a single heart replaces the string of rudimentary hearts constituting the dorsal blood-vessel in the lower _Annulosa_, (reaching in one species to the number of one hundred and sixty), is a truth as much needing to be comprised in the history of evolution, as is the formation of a respiratory surface by a branched expansion of the skin. A right conception of the genesis of a vertebral column, includes not only the differentiations from which result the _chorda dorsalis_ and the vertebral segments imbedded in it; but quite as much it includes the coalescence of numerous vertebral processes with their respective vertebral bodies. The changes in virtue of which several things become one, demand recognition equally with those in virtue of which one thing becomes several. Evidently, then, the current statement which ascribes the developmental progress to differentiations alone, is incomplete. Adequately to express the facts, we must say that the transition from the homogeneous to the heterogeneous is carried on by differentiations and accompanying integrations. It may not be amiss here to ask--What is the meaning of these integrations? The evidence seems to show that they are in some way dependent on community of function. The eight segments which coalesce to make the head of a centipede, jointly protect the cephalic ganglion, and afford a solid fulcrum for the jaws, &c. The many bones which unite to form a vertebral skull have like uses. In the consolidation of the several pieces which constitute a mammalian pelvis, and in the anchylosis of from ten to nineteen vertebrae in the sacrum of a bird, we have kindred instances of the integration of parts which transfer the weight of the body to the legs. The more or less extensive fusion of the tibia with the fibula and the radius with the ulna in the ungulated mammals, whose habits require only partial rotations of the limbs, is a fact of like meaning. And all the instances lately given--the concentration of ganglia, the replacement of many pulsating blood-sacs by fewer and finally by one, the fusion of two uteri into a single uterus--have the same implication. Whether, as in some cases, the integration is merely a consequence of the growth which eventually brings into contact adjacent parts performing similar duties; or whether, as in other cases, there is an a
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