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enera, families and orders, of grouping of the somites. They are anomotagmic, as well as anomomeristic. [Illustration: FIG. 33.--The alimentary canal and gastric glands of a scorpion (A) and of Limulus (B). From Lankester, "Limulua an Arachnid." ps, Muscular suctorial enlargement of the pharynx. sal, Prosomatic pair of gastric caeca in Scorpio, called salivary glands by some writers. c^1, and c^2, The anterior two pairs of gastric caeca and ducts of the mesosomatic region. c^3, c^4 and c^5. Caeca and ducts of Scorpio not represented in Limulus. M, The Malpighian or renal caecal diverticula of Scorpio. pro, The proctodaeum or portion of gut leading to anus and formed embryologically by an inversion of the epiblast at that orifice.] When it is admitted--as seems to be reasonable--that the primitive Arachnida would, like the primitive Crustacea, be anomomeristic and anomotagmic, we shall not demand of claimants for the rank of primitive Arachnids agreement with Limulus and Scorpio in respect of the exact number of their somites and the exact grouping of those somites; and when we see how diverse are the modifications of the branches of the appendages both in Arachnida and in other classes of Arthropoda (q.v.), we shall not over-estimate a difference in the form of this or that appendage exhibited by the claimant as compared with the higher Arachnids. With those considerations in mind, the claim of the extinct group of the trilobites to be considered as representatives of the lower and more primitive steps in the Arachnidan genealogy must, it seems, receive a favourable judgment. They differ from the Crustacea in that they have only a single pair of prae-oral appendages, the second pair being definitely developed as mandibles. This fact renders their association with the Crustacea impossible, if classification is to be the expression of genetic affinity inferred from structural coincidence. On the contrary, this particular point is one in which they agree with the higher Arachnida. But little is known of the structure of these extinct animals; we are therefore compelled to deal with such special points of resemblance and difference as their remains still exhibit. They had lateral eyes[5] which resemble no known eyes so closely as the lateral eyes of Limulus. The general form and structure of their prosomatic carapace are in many striking features identical with that of Limulus. The tril
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