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an eighteen and may be grouped only as head (prosoma) and trunk or may be further differentiated. A telsonic tergal shield of greater or less size is always present, which may be imperfectly divided into well-marked but immovable tergites indicating incompletely differentiated somites. The single pair of palpiform appendages in front of the mouth has been found in one instance to be antenniform, whilst the numerous post-oral appendages in the same genus were bi-ramose. The position of the genital apertures is not known. Compound lateral eyes present; median eyes wanting. The body and head have the two pleural regions of each somite flattened and expanded on either side of the true gut-holding body-axis. Hence the name of the sub-class signifying tri-lobed, a condition realized also in the Xiphosurous Arachnids. The members of this group, whilst resembling the lower Crustacea (as all lower groups of a branching genealogical tree must do), differ from them essentially in that the head exhibits only one prosthomere (in addition to the eye-bearing prosthomere) with palpiform appendages (as in all Arachnida) instead of two. The Anomomeristic Arachnida form a single sub-class, of which only imperfect fossil remains are known. Sub-class (of the Anomomeristica). TRILOBITAE.--The single sub-class Trilobitae constitutes the grade Anomomeristica. It has been variously divided into orders by a number of writers. The greater or less evolution and specialization of the metasomatic carapace appears to be the most important basis for classification--but this has not been made use of in the latest attempts at drawing up a system of the Trilobites. The form of the middle and lateral regions of the prosomatic shield has been used, and an excessive importance attached to the demarcation of certain areas in that structure. Sutures are stated to mark off some of these pieces, but in the proper sense of that term as applied to the skeletal structures of the Vertebrata, no sutures exist in the chitinous cuticle of Arthropoda. That any partial fusion of originally distinct chitinous plates takes place in the cephalic shield of Trilobites, comparable to the partial fusion of bony pieces by suture in Vertebrata, is a suggestion contrary to fact. The Trilobites are known only as fossils, mostly Silurian and prae-Silurian; a few are found in Carboniferous and Permian strata. As
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