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in a situation which is EXCENTRIC in the nervous system--that is, distant from the nervous centres. This mode of action has not, I think, been hitherto distinctly understood by physiologists. "Many of the phenomena of this principle of action, as they occur in the limbs, have certainly been observed. But, in the first place, this function is by no means confined to the limbs; for, while it imparts to each muscle its appropriate tone, and to each system of muscles its appropriate equilibrium or balance, it performs the still more important office of presiding over the orifices and terminations of each of the internal canals in the animal economy, giving them their due form and action; and, in the second place, in the instances in which the phenomena of this function have been noticed, they have been confounded, as I have stated, with those of sensation and volition; or, if they have been distinguished from these, they have been too indefinitely denominated instinctive, or automatic. I have been compelled, therefore, to adopt some new designation for them, and I shall now give the reasons for my choice of that which is given in the title of this paper--'Reflex Functions.' "This property is characterized by being EXCITED in its action and REFLEX in its course: in every instance in which it is exerted an impression made upon the extremities of certain nerves is conveyed to the medulla oblongata or the medulla spinalis, and is reflected along the nerves to parts adjacent to, or remote from, that which has received the impression. "It is by this reflex character that the function to which I have alluded is to be distinguished from every other. There are, in the animal economy, four modes of muscular action, of muscular contraction. The first is that designated VOLUNTARY: volition, originated in the cerebrum and spontaneous in its acts, extends its influence along the spinal marrow and the motor nerves in a DIRECT LINE to the voluntary muscles. The SECOND is that of RESPIRATION: like volition, the motive influence in respiration passes in a DIRECT LINE from one point of the nervous system to certain muscles; but as voluntary motion seems to originate in the cerebrum, so the respiratory motions originate in the medulla oblongata: like the voluntary motions, the motions of respirations are spontaneous; they continue, at least, after the eighth pair of nerves have been divided. The THIRD kind of muscular action in the anima
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