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eserved entire. "All the kinds of muscular motion may be unduly excited, but the reflex function is peculiar in being excitable in two modes of action, not previously subsisting in the animal economy, as in the case of sneezing, coughing, vomiting, etc. The reflex function also admits of being permanently diminished or augmented and of taking on some other morbid forms, of which I shall treat hereafter. "Before I proceed to the details of the experiments upon which this disposition rests, it may be well to point out several instances in illustration of the various sources of and the modes of muscular action which have been enumerated. None can be more familiar than the act of swallowing. Yet how complicated is the act! The apprehension of the food by the teeth and tongue, etc., is voluntary, and cannot, therefore, take place in an animal from which the cerebrum is removed. The transition of food over the glottis and along the middle and lower part of the pharynx depends upon the reflex action: it can take place in animals from which the cerebrum has been removed or the ninth pair of nerves divided; but it requires the connection with the medulla oblongata to be preserved entirely; and the actual contact of some substance which may act as a stimulus: it is attended by the accurate closure of the glottis and by the contraction of the pharynx. The completion of the act of deglutition is dependent upon the stimulus immediately impressed upon the muscular fibre of the oesophagus, and is the result of excited irritability. "However plain these observations may have made the fact that there is a function of the nervous muscular system distinct from sensation, from the voluntary and respiratory motions, and from irritability, it is right, in every such inquiry as the present, that the statements and reasonings should be made with the experiment, as it were, actually before us. It has already been remarked that the voluntary and respiratory motions are spontaneous, not necessarily requiring the agency of a stimulus. If, then, an animal can be placed in such circumstances that such motions will certainly not take place, the power of moving remaining, it may be concluded that volition and the motive influence of respiration are annihilated. Now this is effected by removing the cerebrum and the medulla oblongata. These facts are fully proved by the experiments of Legallois and M. Flourens, and by several which I proceed to det
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