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There are nine bones in the plastron of the tortoise. "The conclusion to be drawn from this is that every sternum, provided that it is not inhibited in its development by some obstacle, is composed of _nine elementary parts_" (p. 105). These nine bones are in Geoffroy's nomenclature, the episternals, the hyosternals, the hyposternals, the xiphisternals, which are all paired bones, and the entosternal, which is unpaired. The arrangement of them is in the tortoise:-- Episternal---------------------------Episternal |\__ __/| | \__ __/ | | \__ __/ | | \__ Entosternal __/ | | __/ \__ | | __/ \__ | | __/ \__ | |/ \| Hyosternal Hyosternal | | | | | | | | Hyposternal-------------------------Hyposternal | | | | | | | | Xiphisternal------------------------Xiphisternal. The articulations in the tortoise are indicated by the connecting lines. Geoffroy tries to show that the sternum in other animals is composed of these nine bones, or at least of a certain number of them, always in the same invariable relative positions. Thus in birds the sternum consists of five pieces, of a huge keeled entosternal, and of two "annexes" on either side, which are the hyo-and hyposternals. These are separate only in young birds. Occasionally, especially in young birds, rudiments of episternals and xiphisternals also occur. The minuteness of the episternals and the xiphisternals may be attributed to the gigantic size of the entosternal, in accordance with the _Loi de balancement_. In the other air-breathing Vertebrates the nine sternal elements can according to Geoffroy be discovered without great difficulty. But when we come to the determination of the sternum in fishes, difficulties abound, which Geoffroy solves in the following way. He points out that between the clavicles (_cleithra_) and the hyoid bone (_basihyal_) in fishes there is a long median bo
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