ht of these subordinate
hypotheses are required, namely, that--
"1. The emission of the gemmules takes place, or may take place in all
states of the cell.
"2. The quantity of gemmules emitted from every cell is very great.
"3. The minuteness of the gemmules is extreme.
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"4. The gemmules possess two sorts of affinity, one of which might be
called _propagative_, and the other _germinative_ affinity.
"5. By means of the propagative affinity all the gemmules emitted by all
the cells of the individual flow together and become condensed in the cells
which compose the sexual organs, whether male or female (embryonal vesicle,
cells of the embryo, pollen grains, fovilla, antherozoids, spermatozoids),
and likewise flow together and become condensed in the cells which
constitute the organs of a sexual or agamic reproduction (buds, spores,
bulbilli, portions of the body separated by scission, &c.).
"6. By means of the germinative affinity, every gemmule (except in cases of
anomalies or monstrosities) can be developed only in cells homologous with
the mother-cells of the cell from which they originated. In other words,
the gemmules from any cell can only be developed in unison with the cell
preceding it in due order of succession, and whilst in a nascent state.
"7. Of each kind of gernmule a great number perishes; a great number
remains in a dormant state through many generations in the bodies of
descendants; the remainder germinate and reproduce the mother-cell.
"8. Every gemmule may multiply itself by a process of scission into any
number of equivalent gemmules."
Mr. Darwin has published a short notice in reply to Professor Delpino, in
_Scientific Opinion_ of October 20, 1869, p. 426. In this reply he admits
the justice of Professor Delpino's attack, but objects to the alleged
necessity of the first subordinate hypothesis, namely, that the emission of
gemmules takes place in all states of the cell. But if this is not the
case, then a great part of the utility and distinction of pangenesis is
destroyed, or as Mr. Lewes justly says,[225] "If gemmules produce whole
cells, we have the very power which was pronounced mysterious in larger
organisms."
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Mr. Darwin also does not see the force of the objection to the power of
self-division which must be asserted of the gemm
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