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soderm. The Coelentera may thus be briefly defined as Metazoa which exhibit two embryonic cell-layers only,--the ectoderm and endoderm,--their body-cavities being referable to a single cavity or coelenteron in the endoderm. Their position in the animal kingdom and their main subdivisions may be expressed in the following table:-- I. PROTOZOA. II. PARAZOA or PORIFERA. III. METAZOA. | +----------+--------------+ | | Coelentera Triploblastica = Diploblastica. (including Coelomata). | +------+-----------+-----------------+ | | | Hydromedusae. Scyphozoa. Ctenophora. | +--------+----------+ | | Scyphomedusae. Anthozoa. In the above-given classification, the Scyphomedusae, formerly included with the Hydromedusae as Hydrozoa, are placed nearer the Anthozoa. The reasons for this may be stated briefly. The HYDROMEDUSAE are distinguished from the Scyphozoa chiefly by negative characters; they have no stomodaeum, that is, no ingrowth of ectoderm at the mouth to form an oesophagus; they have no mesenteries (radiating partitions) which incompletely subdivide the coelenteron; and they have no concentration of digestive cells into special organs. Their ectodermal muscles are mainly longitudinal, their endodermal muscles are circularly arranged on the body-wall. Their sexual cells are (probably in all cases) produced from the ectoderm, and lie in those radii which are first accentuated in development. They typically present two structural forms, the non-sexual hydroid and the sexual medusoid; in such a case there is an alternation of generations (metagenesis), the hydroid giving rise to the medusoid by a sexual gemmation, the medusoid bearing sexual cells which develop into a hydroid. In some other cases medusoid develops directly from medusoid (hypogenesis), whether by sexual cells or by gemmation. The medusoids have a muscular velum of ectoderm and mesogloea only. The SCYPHOZOA have the following features in common:--They typically exhibit an ectodermal stomodaeum; partitions or mesenteries project into their coelenteron from the body-wall, and on these are generally concentrated digestive cells (to f
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