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so hasten the elimination of disused and useless parts. DROOPING EARS AND DETERIORATED INSTINCTS. Mr. Galton has pointed out that existing races and existing organs are only kept at their present high pitch of organic excellence by the stringent and incessant action of natural or artificial selection; and the simple relaxation or withdrawal of such selective influences will almost necessarily result in a certain amount of deterioration, independently even of the principle of economy.[21] I think that this cessation of a previous selective process will account for the drooping--but _not diminished_--ears of various domesticated animals (human preference and increased weight evidently aiding), and also for the inferior instincts seen in them and in artificially-fed caterpillars of the silk-moth, which now "often commit the strange mistake of devouring the base of the leaf on which they are feeding, and consequently fall down." Anyhow, I fail to see that anything is proved by this latter case, except that natural instinct may be perverted or aborted under unnatural conditions and a changed method of selection which abolishes the powerful corrective formerly supplied by natural selection. WINGS AND LEGS OF DUCKS AND FOWLS. The reduced wings and enlarged legs of domesticated ducks and fowls are attributed by Darwin and Spencer to the inheritance of the effects of use and disuse. But the inference by no means follows. Natural selection would usually favour these adaptive changes, and they would also have been aided by an artificial selection which is often unconscious or indirect. Birds with diminished power of flight would be less difficult to keep and manage, and in preserving and multiplying such birds man would be unconsciously bringing about structural changes which would easily be regarded as effects of use and disuse. "About eighteen centuries ago Columella and Varro speak of the necessity of keeping ducks in netted enclosures like other wild fowl, so that at this period there was danger of their flying away."[22] Is it not probable that the best fliers would escape most frequently, or would pine most if kept confined? On the other hand, birds with lessened powers of flight would not be eliminated as under natural conditions, but would be favoured; and natural selection, together with artificial selection of the most flourishing birds, would thicken and strengthen the legs to meet increased demands upon t
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