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of the transformation of species. Against these cases, and against using the palaeontological archives as a basis for the construction of genealogical trees in general, the weighty and apparently decisive objection is urged, that nowhere are the soft parts of the earlier forms of life preserved, and that it is impossible to establish relationships with any certainty on the basis of hard parts only, such as bones, teeth and shells. Even Haeckel admits that snails of very different bodily structure may form very similar and even hardly distinguishable shells. Fleischmann further asserts that Haeckel's "fundamental biogenetic law" has utterly collapsed. "Recapitulation" does not occur. Selenka's figures of ovum-segmentation show that there are specific differences in the individual groups. The origin and development of the blastoderm or germinal disc has nothing to do with recapitulation of the phylogeny. It is not the case that the embryos of higher vertebrates are indistinguishable from one another. Even the egg-cell has a specific character, and is totally different from any unicellular organism at the Protistan level. The much-cited "gill-clefts" of higher vertebrates in the embryonic stage are not persistent reminiscences of earlier lower stages; they are rudiments or primordia shared by all vertebrates, and developing differently at the different levels; (thus in fishes they become breathing organs, and in the higher vertebrates they become in part associated with the organs of hearing, or in part disappear again). Though Fleischmann's vigorous protest against over-hastiness in construction and over-confidence on the part of the adherents of the doctrine of descent is very interesting, and may often be justified in detail, it is difficult to resist the impression that the wheat has been rejected with the chaff.(23) Even a layman may raise the following objections: Admitting that the great groups of forms cannot be traced back to one another, the palaeontological record still proves, though it may be only in general outline, that within each phylum there has been a gradual succession and ascent of forms. How is the origin of what is new to be accounted for? Without doing violence to our thinking, without a sort of intellectual autonomy, we cannot rest content with the mere fact that new elements occur. So, in spite of all "difficulties," the assumption of _an actual descent_ quietly forces itself upon us as the
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