hromosomes united end to end in synapsis and
separated in the first maturation mitosis, which is therefore
reductional. The odd chromosome and the _m_-chromosomes show no
longitudinal split in these figures, but they may appear as in figure
249. Occasionally one of the tetrads takes the form of a cross (fig.
249). In this figure the split "accessory" (_x_) lies against the
nuclear membrane and the archoplasmic material for the spindle is seen
along one side of the nucleus. It is certain here that the spindle
fibers come from extranuclear material, not from nuclear substance, as
Paulmier ('99) describes for _Anasa tristis_.

Figures 250 and 251 show the first maturation mitosis as it usually
appears in sections from mercuro-nitric material stained with
iron-haematoxylin. The odd chromosome is always more or less eccentric
and is attached by a spindle fiber to one pole. In Hermann material,
considerably destained, the tetrads and the odd chromosome appear as in
figures 252, 253, and 254, the tetrads being in position for a
transverse division. The odd chromosome is always so placed that its
longitudinal split is at right angles to the axis of the spindle, as
though it were to divide in this mitosis. It does not do so, however,
but goes to one daughter cell, always lagging behind, as is shown in
figures 255 and 256. Figures 257, _a_ and _b_, are polar plates of the
first mitosis with 11 and 12 chromosomes, respectively, and figures 258,
_a_, _b_, and _c_, show the polar plates (_a_ and _c_) each containing
11 chromosomes, and the odd chromosome at a different level (_b_). The
latter is a view of the anaphase which one often gets at three foci in
one section. Figures 259, _a_ and _b_, are equatorial plates of the
second mitosis with 11 and 12 chromosomes respectively. Figure 260 shows
a side view of the second spindle in metaphase, and figure 261 in
anaphase. Figures 262 and 263 are daughter plates from two spindles
showing the chromosome content of the two equal classes of spermatozoa,
one class containing 11 ordinary chromosomes, the other 11 ordinary
chromosomes plus the odd heterochromosome, for the odd chromosome
divides with the others in the second spindle as in Orthoptera (McClung
and Sutton).

In figures 264 and 265 (plate XV) are seen the telophase of the two
kinds of second spermatocytes, one (fig. 265) showing the divided odd
chromosome, which continues to stain more deeply after the others have
become diffuse