toes
about two-thirds webbed (Fig. 1d).

Skin of venter coarsely areolate; skin of flanks, throat,
chest, undersurfaces of arms, tibia, tarsi, dorsal surfaces
of thighs, tarsi, hands, and feet smooth; skin of dorsal
surfaces of tibia, forearm, back, and top and sides of head
having large horny pustules (sharply spinous in male).

[Illustration: FIG. 1. _Allophryne ruthveni_, male (KU 69890);
(_a_) Dorsum. (_b_) Thenar view of right hand. (_c_) Lateral profile of
head. (_d_) Plantar view of right foot. x 3.5.]

_Color._--Dorsum gray with irregular network of black lines
and elongate blotches; flanks and labial region black with
large white ocelli; dorsal surfaces of limbs gray, marked as
follows: two large, elongate white spots on each thigh,
concealed white spot on base of upper arm, black-edged gray
transverse bars on forearms and shanks, white spot on each
knee and elbow; ventral surfaces pale gray; black-edged
white spot on ventral surface of thigh on each side of anal
opening; chin and throat dark gray with white spots; vocal
sac in male black (Fig. 1a and c).

Gaige (1926) briefly described the color, which conforms to
the above in all particulars. The paratype (MCZ 11790) has
lost the gray color after 40 years in preservation; now
(1966) the ground-color is cream-brown, and the dorsal
spotting, noted by Gaige as being black, is now brown.

The spots on the feet, tarsi, knees, thighs, flanks and
upper arm are white in preservative, but in life possibly
were red or yellow. These colors usually fade to white in
preservative. Red or yellow spots are common aposematic
colors in frogs.

_Variation._--Eight measurements were taken on each specimen
and four ratios were computed; these are summarized in Table
1. Gaige's illustration of the holotype shows that it has a
greatly reduced pattern, whereas the paratype and three of
the other five known specimens have relatively large and
numerous spots. The male (KU 69890) and one female (AMNH
70108) have a reduced pattern intermediate between that of
the holotype and the four other specimens.

TABLE I.--Variation in Measurements and Proportions of Allophryne
ruthveni. (Ranges in parentheses below means.)

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Character | Male (1) |