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sustain itself. Haberlandt furthermore records a phenomenon perhaps analogous to this on the grape-vine, but with this exception the case described by him is unique. In order to pass any further judgment regarding it, we should have to ascertain whether the whole phenomenon is not a case of so-called adaptation; if so, processes should be found in nature, analogous to the poisoning of the hydathodes in this experiment, which result in the destruction of the hydathodes so that in consequence the plant would have gained the power of making good the loss, by means of the substitute organs. Such processes, however, (even through poisoning or through parasites) would be very highly improbable. Equally incredible is the alternative possibility that the new organs would be produced by the plant not as a substitute but as a supplementary apparatus when the old ones would not suffice for secretion in case of very large absorption of water. This also must doubtlessly be rejected, as Haberlandt has observed. Powers of adaptation should, of course, according to Darwinism, be gradually acquired in the struggle for existence, as in that case they should also have stability; but since this is not possessed by the new organs, the presumption is that they do not possess the character of adaptation. They are therefore new organs that originated after an entirely unnatural and unforeseen interference with the normal vital functions and in consequence of a self-regulating activity of the organism. What then is there in the whole phenomenon worthy of notice with regard to the theory of Descent? 1. An immediately well adapted new organ has here originated very suddenly without any previous incipient formation, without gradual perfection and without stages of transition. 2. In its formation struggle for existence and natural selection are entirely excluded, neither can find any application whatever even according to the newer exposition of Weismann. Haberlandt himself draws this conclusion. 3. If this phenomenon of a suddenly appearing change can take place in the course of the development of the individual, there can be no obvious reason why it should not take place in the same manner (without natural selection or struggle for existence) in the course of the phylogenetic development. It is manifestly of the greatest importance that in this case a direct, experimental proof has been given that an organ has originated suddenly
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