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into the developing wing. In C, _cu'_ represents the new cuticle forming beneath the old one, and (_p_) the pouch within which the wing-disc (_w_) lies. Highly magnified. After Gonin, _Bull. Soc. Vaud._ XXX.] As mentioned above, this hidden growth of the wing-rudiments, in butterflies, beetles, flies, bees, and the great majority of the winged insects, has been emphasised by Sharp (1899) as a character contrasting markedly with the outward and visible growth of the wing-rudiments in such insects as cockroaches, bugs, and dragon-flies. The divergence between the two modes of development is certainly very striking, and a conceivable method of transition from the one to the other is not easy to explain. Sharp has expressed the divergence by the terms _Endopterygota_, applied to all the orders of insects with hidden wing-rudiments (the 'Metabola' or 'Holometabola' of most classifications) and _Exopterygota_, including all those insects whose wing-rudiments are visible throughout growth ('Hemimetabola' and 'Ametabola'). Those curious lowly insects, belonging to the two orders of the Collembola and Thysanura, none of whose members ever develop wings at all, form a third sub-class, the _Apterygota_ (see Classificatory Table, p. 122). Not the wings only, but other structures of the imago, varying in extent in different orders, are formed from the imaginal discs. For example, de Reaumur and G. Newport (1839) found that if the thoracic leg of a late-stage caterpillar were cut off, the corresponding leg of the resulting butterfly would still be developed, although in a truncated condition. Gonin has shown that in the Cabbage White butterfly (_Pieris brassicae_) the legs of the imago are represented, through the greater part of larval life, only by small groups of cells situated within the bases of the larval legs. After the third moult these imaginal discs grow rapidly and the proximal portion of each, destined to develop into the thigh and shin of the butterfly's leg, sinks into a depression at the side of the thorax, while the tip of the shin and the five-segmented foot project into the cavity of the larval leg. Hence we understand that the amputation of the latter by the old naturalists truncated only and did not destroy the imaginal limb. In the blow-fly maggot, Weismann, B.T. Lowne (1890) and J. Van Rees (1888) have shown that the imaginal discs of the legs (fig. 11--1, 2, 3) grow out from deep dermal inpushings. Simple
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