from
Nicaragua. Guenther (1901:267) placed _boulengeri_ in the genus _Hyla_,
and stated that Cope possibly placed _boulengeri_ in the genus
_Scytopis_ on the supposition that it had an accumulation of
"sebaceous glands" above the tympanum. Noble (1918:339) redescribed
_Hyla boulengeri_ on the basis of three specimens from Zelaya
Province, Nicaragua, and noted that the glands were not prominent in
any of the specimens. Duellman (1956:8) showed that _Scytopis hebes_
(generotype of _Scytopis_ by monotypy) is a Phrynohyas, and thus
placed _Scytopis_ Cope, 1862, in the synonymy of _Phrynohyas_
Fitzinger, 1843.

Dunn and Emlen (1932:25) placed _Hyla lancasteri_ Barbour in the
synonymy of _Hyla boulengeri_; the former was known solely from one
juvenile. They made no qualifying statements, but probably they were
impressed by the strongly barred thighs, a coloration known among
Central American hylids at that time only in _Hyla boulengeri_
(Duellman, 1966a:271). Taylor (1952:856) followed Dunn and Emlen with
reservation and noted some differences. Duellman (1966a:271) showed
that the holotype of _lancasteri_ was a juvenile of a species
subsequently named as _Hyla moraviaensis_ by Taylor (1952:865).

In Central America, _Hyla boulengeri_ can be confused only with _Hyla
foliamorta;_ the latter is restricted to central and eastern Panama
and northern Colombia. The snout of _foliamorta_ is more pointed and
protruding, and the vocal sac is darker than in _boulengeri_; the
groin of _foliamorta_ usually is creamy white, whereas _boulengeri_
usually has a dark spot. The skulls differ in that _boulengeri_ has a
frontoparietal fontanelle, the prevomer is larger and elongate,
anteriorly connected to the premaxillary, and posteriorly separated
from the sphenethmoid by cartilage; _foliamorta_ lacks a fontanelle,
the prevomer is smaller, anteriorly separated from the premaxillary by
cartilage, but connected by a bony suture to the sphenethmoid. The
mating call of _boulengeri_ differs by having shorter notes, twice as
many pulses per second, a higher fundamental frequency, and more
closely approximated major frequencies than does that of _foliamorta_.

_Hyla boulengeri_ need not be compared in detail with the other Central
American members of the _Hyla rubra_ group, because all of them are
smaller and have shorter snouts, smoother skin, and dissimilar color
patterns.

_Distribution._--In Central America _Hyla boulengeri_ inhabits the
fo