-vent length and the ratio of the
diameter of the tympanum to that of the eye, have been observed in
this species. The largest individuals are from the Golfo Dulce region
(samples from Piedras Blancas and Rincon de Osa), Puntarenas Province,
Costa Rica. The smallest individuals are from El Recreo, Zelaya
Province, Nicaragua, and from the Caribbean lowlands of Costa Rica.

The diameter of the tympanum is proportionately larger (relative to
the size of the eye) in males from Tilaran, Guanacaste Province; the
tympanum is nearly as large in males from Piedras Blancas, Puntarenas
Province, and Puerto Viejo, Heredia Province, Costa Rica. The lowest
ratios occur in individuals from Almirante, Bocas del Toro, Panama, in
specimens from the Caribbean lowlands of Costa Rica (except Puerto
Viejo), and in those from El Recreo, Zelaya Province, Nicaragua. In
general, the tympanum is proportionately larger in females than in
males; the tympanum is largest in females from the Pacific lowlands
of Costa Rica (Table 5).

Color variation has been observed in individuals from the same
population, as well as in individuals from different localities,
between males and females, and from night to day. In life, most
individuals from the Pacific lowlands of Costa Rica are dark tan to
greenish gray above with dark brown longitudinal stripes that are
entire or broken, but some specimens (mostly males) are dusky brown
and lack longitudinal stripes or an interorbital triangle; females
usually have the dark interorbital triangle and the stripes on the
dorsum. Individuals from Turrialba, Cartago Province, Costa Rica, are
pale olive-tan with olive-brown markings. Individuals from Puerto
Viejo, Heredia Province, Costa Rica, are uniformly yellowish brown
with or without dark longitudinal stripes. Specimens from El Recreo,
Zelaya Province, Nicaragua, are like those from Puerto Viejo. Males
from Almirante, Bocas del Toro, Panama, are pale brown with dark brown
longitudinal stripes and an indistinct interorbital triangle. Females
have a distinct interorbital triangle and dark brown blotches on the
thighs and shanks.

By night, the dorsum usually is pale yellow, and the belly is creamy
white. By day, the dorsum is dark tan; the stripes and spots are
darker, and the belly is yellowish white. Taylor (1952) noticed that
considerable variation in color pattern occurred from night to day in
individuals from Turrialba, Cartago Province, Costa Rica. At night
som