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rst levels taken together constitute the so-called "voluntary circuit" (see Fig. 2). In addition to this general demarcation of functions as higher and lower--first, second, and third level--in their anatomical seat, many interesting discoveries have been made in the localization of the simpler functions in the cortex itself. The accompanying figures (Figs. 3 and 4) will show the principle centres which have been determined; and it is not necessary to dwell upon additional details which are still under discussion. The areas marked out are in general the same on both hemispheres, and that is to say that most of the centres are duplicated. The speech centres, however, are on one side only. And in certain cases the nervous fibres which connect the cortex with the body-organs cross below the brain to the opposite side of the body. This is always true in cases of muscular movement; the movements of the right side of the body are controlled by the left hemisphere, and _vice versa_. The stimulations coming in from the body to the brain generally travel on the same side, although in certain cases parallel impulses are also sent over to the other hemisphere as well. For example, the very important optic nerve, which is necessary to vision, comes from each eye separately in a large bunch of fibres, and divides at the base of the brain, so that each eye sends impulses directly to the visual centres of both hemispheres. [Illustration: FIG. 3.--Outer surface of left hemisphere of the brain (modified from Exner): _a_, fissure of Rolando; _b_, fissure of Sylvius.] [Illustration: FIG. 4.--Inner (mesial) surface of the right hemisphere of the brain (modified from Schaefer and Horsley). In both figures the shaded area is the motor zone.] Of all the special questions which have arisen about the localization of functions in the nervous system, that of the function of certain areas known as "motor centres" has been eagerly discussed. The region on both sides of the fissure of Rolando in Fig. 3 contains a number of areas which give, when stimulated with electricity, very definite and regular movements of certain muscles on the opposite side of the body. By careful exploration of these areas the principal muscular combinations--those for facial movements, neck movements, movements of the arm, trunk, legs, tail, etc.--have been very precisely ascertained. It was concluded from these facts that these areas were respectively the centres
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