d rubbed against the insect's body. In
several closely allied genera, as in Dielytra, &c., there are two
perfect nectaries, the pistil is straight, and the hood slips off on
either side, according as the bee sucks either nectary. Now, I have
examined several flowers of _Corydalis tuberosa_, in which both
nectaries were equally developed and contained nectar; in this we see
only the redevelopment of a partially aborted organ; but with this
redevelopment the pistil becomes straight, and the hood slips off in
either direction; so that these flowers have acquired the perfect
structure, so well adapted for insect agency, of Dielytra and its
allies. We cannot attribute these coadapted modifications to chance, or
to correlated variability; we must attribute them to reversion to a
primordial condition of the species.
The peloric flowers of Pelargonium have their five petals in all
respects alike, and there is no nectary; so that they resemble the
symmetrical flowers of the closely allied Geranium-genus; but the
alternate stamens are also sometimes destitute of anthers, the
shortened filaments being left as rudiments, and in this respect they
resemble the symmetrical flowers of the closely allied genus, Erodium.
Hence we are led to look at the peloric flowers of Pelargonium as
having probably reverted to the state of some primordial form, the
progenitor of the three closely related genera of Pelargonium,
Geranium, and Erodium.
In the peloric form of _Antirrhinum majus_, appropriately called the
"_Wonder_," the tubular and elongated flowers differ wonderfully from
those of the common snapdragon; the calyx and the mouth of the corolla
consist of six equal lobes, and include six equal instead of four
unequal stamens. One of the two additional stamens is manifestly formed
by the development of a microscopically minute papilla, which may be
found at the base of the upper lip of the flower in all common
snapdragons, at least in nineteen plants examined by me. That this
papilla is a rudiment of a stamen was well shown by its various degrees
of development in crossed plants between the common and peloric
Antirrhinum. Again, a peloric _Galeobdolon luteum_, growing in my
garden, had five equal petals, all striped like the ordinary lower lip,
and included five equal instead of four unequal stamens; b
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