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umination, is a voluntary eructation. "The pointed knots, like little papillae, in the stomachs of ruminating beasts, are also of great use, namely, for the tasting of the meat. The inner membrane of the first three venters is fibrous (like the gustatory papillae of the tongue) and not glandulous; the fourth only being glandulous, as in a man. Of the fibres of this membrane, and the nervous, are composed those pointed knots, which are, both in substance and shape, altogether like to those upon the tongue. Whence I doubt not, but that the said three ventricles, as they have a power of voluntary motion, so, likewise, that they are the seat of taste, and as truly the organs of that sense, as is the tongue itself." [Illustration: Skull of Domestic Ox, from a specimen in the Royal College of Surgeons.] The mouth of animals of the Ox Tribe contains, when full, thirty-two teeth. Six molars in each jaw, above, below, and on either side; and eight incisors in the lower jaw. In the upper jaw there are no incisors; but instead thereof a fibrous and elastic pad, or cushion, which covers the convex extremity of the anterior maxillary bone, and which is well worthy of observation. The final cause of this pad (which stands in the place of upper incisor teeth) and the part it plays in the procuring of food, is thus described by Youatt. "The grass is collected and rolled together by means of the long and moveable tongue; it is firmly held between the lower cutting teeth and the pad, the cartilaginous upper lip assisting in this; and then by a sudden nodding motion of the head, the little roll of herbage is either torn or cut off, or partly both torn and cut. "The intention of this singular method of gathering the food, it is somewhat difficult satisfactorily to explain. It is peculiar to ruminants, who have one large stomach, in which the food is kept as a kind of reservoir until it is ready for the action of the other stomachs. While it is kept there it is in a state of maceration; it is exposed to the united influence of moisture and warmth, and the consequence of this is, that a species of decomposition sometimes commences, and a vast deal of gas is extricated. "That this should not take place in the natural process of retention and maceration, nature possibly established this mechanism for the first gathering of the food. It is impossible that half of that which is thus procured can be fairly cut through; part will be to
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