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ce. For purposes of reference, a standard of comparison is required; and this standard, so long as its nature is not overlooked, may, indeed must be, to some extent, an arbitrary one. Thus in the phanerogamous plants there is assumed to exist, in all cases, an axis (stem, branches, roots, thalamus, &c.), bearing leaves and flowers. These latter consist of four whorls, calyx, corolla, stamens, and pistils, each whorl consisting of so many separate pieces in determinate position and numbers, and of regular proportionate size. A very close approach to such a flower occurs normally in _Limnanthes_ and _Crassula_, and, indeed, in a large proportion of all flowers in an early stage of development. To a standard type, such as just mentioned, all the varied forms that are met with, either in normal or abnormal morphology, may be referred by bearing in mind the different modifications and adaptations that the organs have to undergo in the course of their development. Some parts after a time may cease to grow, others may grow in an inordinate degree, and so on; and thus, great as may be the ultimate divergences from the assumed standard, they may all readily be explained by the operation, simply or conjointly, of some of the four principal causes of malformation before alluded to. The fact that so many and such varied changes can thus readily be explained is not only a matter of convenience, but may be taken as evidence that the standard of reference is not wholly arbitrary and artificial, but that it is a close approximation to the truth. It has already been said that an arrangement like that here considered as typical is natural to some flowers in their adult state, and to a vast number in their immature condition. It would be no extravagant hypothesis to surmise that this was the primitive structure of the flower in the higher plants. Variations from it may have arisen in course of time, owing to the action of an inherent tendency to vary, or from external circumstances and varied requirements which may have induced corresponding adaptations, and which may have been transmitted in accordance with the principle of hereditary transmission. This hypothesis necessarily implies a prior simplicity of organisation, of which, indeed, there is sufficient proof; many cases of malformation can thus be considered as so many reversions to the ancestral form. Thus, teratology often serves as an aid in the study of morphology in genera
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