FREE BOOKS

Author's List




PREV.   NEXT  
|<   22   23   24   25   26   27   28   29   30   31   32   33   34   35   36   37   38   39   40   41   42   43   44   45   46  
47   48   49   50   51   52   53   54   55   56   57   58   59   60   61   62   63   64   65   66   67   68   69   70   71   >>   >|  
lly coiled in numerous close turns, motile, but apparently owing to flexile movements, as no cilia are found. II. ORDER--TRICHOBACTERINAE. Vegetative body of branched or unbranched cell-filaments, the segments of which separate as swarm-cells (_Gonidia_). 1. _Family_--TRICHOBACTERIACEAE. Characters those of the Order. (a) Filaments rigid, non-motile, sheathed:--_Crenothrix_ (Cohn), filaments unbranched and devoid of sulphur particles; _Thiothrix_ (Winogr.), as before, but with sulphur particles; _Cladothrix_ (Cohn), filaments branched in a pseudo-dichotomous manner. (b) Filaments showing slow pendulous and creeping movements, and with no distinct sheath:--_Beggiatoa_ (Trev.), with sulphur particles. The principal objections to this system are the following:--(1) The extraordinary difficulty in obtaining satisfactory preparations showing the cilia, and the discovery that these motile organs are not formed on all substrata, or are only developed during short periods of activity while the organism is young and vigorous, render this character almost nugatory. For instance, _B. megatherium_ and _B. subtilis_ pass in a few hours after commencement of growth from a motile stage with peritrichous cilia, into one of filamentous growth preceded by casting of the cilia. (2) By far the majority of the described species (over 1000) fall into the three genera--_Micrococcus_ (about 400), _Bacillus_ (about 200) and _Bactridium_ (about 150), so that only a quarter or so of the forms are selected out by the other genera. (3) The monotrichous and lophotrichous conditions are by no means constant even in the motile stage; thus _Pseudomonas rosea_ (Mig.) may have 1, 2 or 3 cilia at either end, and would be distributed by Fischer's classification between _Bactrinium_ and _Bactrillum_, according to which state was observed. In Migula's scheme the attempt is made to avoid some of these difficulties, but others are introduced by his otherwise clever devices for dealing with these puzzling little organisms. The question, What is an individual? has given rise to much difficulty, and around it many of the speculations regarding pleomorphism have centred without useful result. If a tree fall apart into its constituent cells periodically we should have the same difficulty on a larger and more complex scale. The fact that every bacterial cell in a species in most cases app
PREV.   NEXT  
|<   22   23   24   25   26   27   28   29   30   31   32   33   34   35   36   37   38   39   40   41   42   43   44   45   46  
47   48   49   50   51   52   53   54   55   56   57   58   59   60   61   62   63   64   65   66   67   68   69   70   71   >>   >|  



Top keywords:
motile
 

filaments

 

sulphur

 

particles

 
difficulty
 
Filaments
 

showing

 
genera
 

species

 

growth


movements

 

unbranched

 
branched
 

Fischer

 
distributed
 
Bactrillum
 

Bactrinium

 

classification

 
observed
 

difficulties


attempt

 

Migula

 

scheme

 
numerous
 

monotrichous

 
lophotrichous
 

conditions

 

selected

 

quarter

 

constant


coiled

 

introduced

 
Pseudomonas
 

clever

 

constituent

 

periodically

 
result
 
bacterial
 

larger

 

complex


centred

 

pleomorphism

 

organisms

 

question

 
puzzling
 

dealing

 
Bactridium
 

devices

 
individual
 

speculations