rates the two cells of the
anther to some distance.

[Illustration: Fig. 295. Diagram of the lower part of an anther, cut
across above, and the upper part of a leaf, to show how the one answers
to the other; the filament to petiole, the connective to midrib; the two
cells to the right and left halves of the blade.]

289. A simple conception of the morphological relation of an anther to a
leaf is given in Fig. 295, an ideal figure, the lower part representing
a stamen with the top of its anther cut away; the upper, the
corresponding upper part of a leaf.

290. So anthers are generally _two-celled_. But as the pollen begins to
form in two parts of each cell (the anterior and the posterior),
sometimes these two strata are not confluent, and the anther even at
maturity may be _four-celled_, as in Moonseed (Fig. 296); or rather, in
that case (the word _cell_ being used for each lateral half of the
organ), it is _two-celled_, but the cells _bilocellate_.

[Illustration: Fig. 296. Stamen of Moonseed, with anther cut across;
this 4-celled, or rather 4-locellate.]

[Illustration: Fig. 297. Stamen of Pentstemon pubescens; the two
anther-cells diverging, and almost confluent.]

[Illustration: Fig. 298. Stamen of Mallow; the anther supposed to answer
to that of Fig. 297, but the cells completely confluent into one.]

[Illustration: Fig. 299. Stamen of Globe Amaranth; very short filament
bearing a single anther-cell; it is open from top to bottom, showing the
pollen within.]

[Illustration: Fig. 300-305. Stamens of several plants of the Labiate or
Mint Family. Fig. 300. Of a Monarda: the two anther-cells with bases
divergent so that they are transverse to the filament, and their
contiguous tips confluent, so as to form one cell opening by a
continuous line. Fig. 301. Of a Calamintha: the broad connective
separating the two cells. Fig. 302. Of a Sage (Salvia Texana); with long
and slender connective resembling forks of the filament, one bearing a
good anther-cell; the other an abortive or poor one. Fig. 303. Another
Sage (S. coccinea), with connective longer and more thread-shaped, the
lower fork having its anther-cell wholly wanting. Fig. 304. Of a White
Sage, Audibertia grandiflora; the lower fork of connective a mere
vestige. Fig. 305. Of another White Sage (A. stachyoides), the lower
fork of connective suppressed.]

291. But anthers may become _one-celled_, and that either by confluence
or by suppression.

292. By co