ing from nasal septum, and (3) a smooth dorsal
surface on carapace in adult males. _T. m. calvatus_ and _T.
m. muticus_ resemble _T. ferox_ in having a smooth dorsal
surface on carapace in adult males, but differ from _T.
ferox_ in having: (1) no tubercles along anterior edge of
carapace, and (2) no ridge projecting from nasal septum.

_Description of holotype._--Carapace circular, widest at
region of bridge; margin entire; dorsal surface smooth;
anterior margin of carapace lacking tubercles; blunt
vertebral ridge evident anteriorly; maximum length, 53.1 mm;
greatest width, 46.3 mm; greatest depth, 11.5 mm.

Plastron small, extending slightly farther forward than
carapace; anterior lobe truncate with slight midventral
indentation; posterior lobe rounded, sides forming acute
angle; certain features of bony elements of plastron visible
through overlying skin; width of bony bridge, 4.5 mm;
maximum length of plastron, 37.5 mm.

Head extended to level of eyes; head terminating in long,
rounded, flexible snout; nostrils rounded with no ridges
projecting from nasal septum; jaws closed, each covered by
fleshy lips except anteriorly where horny portions of jaws
are exposed; iris with dark stripe through pupil.

Forefeet and hind feet well-webbed and with five digits
each; each limb with nails on first three digits; dorsal
surface of each forelimb with four cornified areas, three of
which have a free edge; each hind limb with two cornified
areas, one smooth on posterodorsal surface and other with
free edge on posteroventral surface.

Tail terminating in flexible point and not extending beyond
posterior edge of carapace; anus to tip of tail, 2.6 mm;
anus to posterior edge of carapace, 8.1 mm.

In preservative: Ground color of carapace dark tan having
pattern of 49 brownish spots; 47 spots circular; two spots
noticeably elongate, one representing fusion of two circular
spots; 17 spots on carapace not exceeding 2.0 mm in
diameter, whereas 32 spots range from 2.5 to 4.0 mm in
diameter; periphery of carapace pale except anteriorly;
maximum width of pale margin (posteriorly), 3.3 mm; junction
of pale margin and dorsal ground color formed by rough-edged
line composed of small, closely-set dots; pattern of fine
punctations and other