radise Park; Muddy Fork of Cowlitz River;
Sunset Park, 5000 ft.; ridge between St. Andrews Park and South Puyallup
River, 6000 ft.; and Owyhigh Lakes, 5350 ft.
Collectively, or individually, where there are as many as six specimens
from a place, the material from Mt. Rainier (Glacier Basin excepted) is
intermediate in color between _T. a. ludibundus_ and _T. a. caurinus_
and no more closely resembles one subspecies than the other. As may be
seen from the cranial measurements recorded above, specimens from Mt.
Rainier, although intermediate between the two subspecies just
mentioned, resemble _ludibundus_ in lesser zygomatic breadth and lesser
cranial breadth (and, it may be added, in lesser dorsolateral inflation
of the braincase), but resemble _caurinus_ in longer skull
(occipitonasal length), longer nasals and greater breadth across the
rows of upper molariform teeth.
In summary: The animals from Mount Rainier, in features of taxonomic
import, are almost exactly intermediate between _T. a. caurinus_ and _T.
a. ludibundus_. Being influenced by considerations of geographic
adjacency, we refer the animals on Mount Rainier to _Tamias amoenus
ludibundus_ (Hollister).
Dalquest's (_op. cit._: 85) explanation of the probable origin of
_Tamias amoenus caurinus_ is pertinent here. He writes: "The chipmunks
of the Olympic Mountains [_caurinus_] probably reached their present
range from the Cascades. Their probable path of emigration was westward
from Mt. Rainier, along the glacial outwash train of Nisqualli Glacier,
to the moraine and outwash apron of the Vashon Glacier and thence to the
Olympics. So similar are the chipmunks of Mt. Rainier and the Olympic
Mountains that Howell (1929) included Mt. Rainier in the range of
_caurinus_."
~Tamias townsendii cooperi~ Baird
Some uncertainty exists concerning the subspecific identity of the
Townsend Chipmunk in southern Washington because Dalquest (Univ. Kansas
Publ. Mus. Nat. Hist., 2:262, April 9, 1948) identified as _Tamias
townsendii cooperi_ specimens that he examined from Yocolt, a place well
within the geographic range of _T. t. townsendii_ as defined by A. H.
Howell (N. Amer. Fauna, 52: fig. 7, p. 107, November 30, 1929). Dalquest
(_op. cit._) referred other specimens, that he did not examine, from Mt.
St. Helens (90654, 231112 and 231114 BS) to _T. t. cooperi_ although
Howell (N. Amer. Fauna, 52:109, November 20, 1929) had previously
identified them as _E. t. townse
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