fold thick, extending to heel; inner metatarsal tubercle small and
elliptical; outer metatarsal tubercle small, flat, and indistinct;
subarticular tubercles small and round; single row of supernumerary
tubercles on proximal segments of each digit; toes moderately short
and slender; length of toes from shortest to longest, 1-2-3-5-4; toes
fully webbed; flap of skin on inner surface of first toe; discs about
same size as those on fingers. Anal opening directed posteroventrally
at middle of thighs; anal sheath moderately elongate; small tubercles
below anal opening. Skin of dorsum rather smooth, somewhat granular
on dorsal surfaces of limbs; skin of chin and belly moderately
granular; that of posterior surfaces of thighs smooth; no thoracic
fold. Tongue nearly round, shallowly notched posteriorly, and free for
about one-fourth its length; vomerine teeth 5-5, situated on rounded
ridges between small inner nares; no vocal slits.

Color (in alcohol) dull olive-green on dorsal surfaces of head, body,
and limbs; flanks dull olive-green with scattered cream-colored spots;
posterior surfaces of thighs grayish brown with faint creamy mottling;
chin gray with cream-colored spots; belly creamy yellow, suffused with
gray posteriorly; undersides of feet and webbing gray; anal stripe
faint, pale cream-color.

_Variation._--The only other known specimen (MNHN 6331) is a female
having a snout-vent length of 53.7 mm. and resembling the specimen
described above in most details of morphology. In MNHN 6331 the
tympanum is completely concealed, and the 8-7 vomerine teeth are
arranged in two irregular rows. The female has more cream-colored
mottling on the flanks and posterior surfaces of the thighs and more
distinct mottling on the throat than the male described above.

_Remarks._--The systematic status of _Cauphias crassus_ Brocchi has
been in doubt since the time of the original description. Brocchi
(1877:130) stated: "Les dernieres phalanges sont obtuses, tronques a
leur extremite anterieure." Brocchi placed the species in his genus
_Cauphias_ (type species, _C. guatemalensis_), which he considered to
be related to _Hylodes_ ( = _Eleutherodactylus_ in the sense used by
Brocchi); he thereby placed _Cauphias_ in his Hylodidae ( =
Leptodactylidae, in part). This idea of relationships was perpetuated
by Barbour (1927:96), who reported on the second known specimen of
_Cauphias guatemalensis_ and stated: "When I dissected the sternum I
was a