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at their unity of construction is due to that fact; but if we go back in thought to the time at which the special peculiarities were introduced which really constituted the order and separated it from other animals, we see that it is by no means clear that it originated with one pair or with one individual, and that, on the contrary, the probabilities are the other way. Although the separation of this order from the rest must have taken place very early, it cannot well have taken place until millions of animals had already come into existence. The prodigality of nature in multiplying animal life is fully acknowledged by Darwin, and that prodigality is apparently greatest in the lowest and most formless type of animal. There being, then, these many millions of living creatures in existence, the external surroundings introduce into them many variations, and among these the special variations to which the vertebrate type is due. It is quite clear that wherever the external surroundings were the same or nearly the same, the variations introduced would be the same or nearly the same. Now, it is far more probable that external surroundings should be the same or nearly the same in many places than that each spot should be absolutely unlike every other spot in these particulars. The beginnings of the vertebrate order would show themselves simultaneously, or at any rate independently, in many places wherever external conditions were sufficiently similar. And the unity of the plan in the vertebrata would be due, not to absolute unity of ancestry, but to unity of external conditions at a particular epoch in the descent of life. Hence it follows that the separation of animals into orders and genera and even into species took place, if not for the most part yet very largely, at a very early period in the history of organic evolution. Of course the descendants of any one of the original vertebrata might, and probably in not a few cases did, branch off into new subdivisions and yet again into further subdivisions, and we are always justified in looking for unity of ancestry among all the species. But it is also quite possible that any species may be regularly descended, without branching off at all, from one of the originals, and that other species that resemble it may owe the resemblance simply to very great similarity of external conditions. To find, for instance, the unity of ancestry between man and the other animals, it will certai
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