al
America and South America are _Eleutherodactylus_ or
_Eupsophus_.
_Distribution._--Low to moderate elevations from Sinaloa,
Mexico, to Guatemala on the Pacific versant; from the
Edwards and Stockton plateaus of Texas to British Honduras
on the Caribbean versant.
_Etymology._--Greek, emendation of _syrrhaptos_, meaning
sewn together in reference to the united outer metatarsals.
Genus ~Tomodactylus~ Guenther, 1900
_Type-species._--_Tomodactylus amulae_ Guenther, 1900.
_Diagnosis and definition._--Small frogs (20 to 35 mm.
snout-vent length) having digital expansions or not, with
transverse groove across tip of each digit; lumbo-inguinal
gland prominently elevated, compact, oval, often patterned;
axillary glands absent; plantar supernumerary tubercles
numerous, more than eight, usually extending between
metatarsal tubercles; tarsus lacking tubercles or folds; toes
free; terminal phalanges T-shaped; sternum cartilaginous,
lacking bony style; sphenethmoid not truncate anteriorly;
nasals in contact medially; maxillary and quadratojugal in
articular contact; anterior arm of squamosal not in contact
with maxillary; dermal cranial elements not involved in
integumentary-cranial co-ossification; prevomers large,
usually bearing dentigerous processes; maxillary and
premaxillary bones dentate; occipital condyles separated;
development direct.
_Composition._--Ten species.
_Distribution._--The southern edge of the Mexican Plateau
from Sinaloa to Veracruuz and onto the Oaxaca highlands and
Sierra Madre del Sur.
_Etymology._--Greek (_tomis_ + _dactylus_) meaning knife
toe; in reference to either the sharp subarticular tubercles
or the unwebbed toes.
DISCUSSION
The preceding definitions only slightly alter the present generic
limits of Mexican leptodactylids. Two species, previously regarded as
_Eleutherodactylus_, are transferred to the new genus _Hylactophryne_.
The arrangement of the species of _Syrrhophus_ and _Tomodactylus_
remains the same as concluded by Dixon (1957), Duellman (1958), and
Firschein (1954) in their reviews of the genera.
Lumbo-inguinal glands are most prominent in the genera _Pleurodema_ and
_Tomodactylus_. Various nondescript glands are present in many genera,
but none is so well developed as those of _Pleurodema_ an
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