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le cells, or asci, mixed with linear thread-like barren cells, called paraphyses, which are regarded by some authors as barren asci. These are placed side by side in juxtaposition with the apex outwards. Each ascus contains a definite number of sporidia, which are sometimes coloured. When mature, the asci explode above, and the sporidia may be seen escaping like a miniature cloud of smoke in the light of the mid-day sun. The disc or surface of the hymenium is often brightly coloured in the genus _Peziza_; tints of orange, red, and brown having the predominance. In _Phacidiacei_, the substance is hard and leathery, intermediate between the fleshy _Elvellacei_ and the more horny of the _Sphaeriacei_. The perithecia are either orbicular or elongated, and the hymenium soon becomes exposed. In some instances, there is a close affinity with the _Elvellacei_, the exposed hymenium being similar in structure, but in all the disc is at first closed. In orbicular forms, the fissure takes place in a stellate manner from the centre, and the teeth are reflexed. In the _Hysteriacei_, where the perithecia are elongated, the fissure takes place throughout their length. As a rule, the sporidia are more elongated, more commonly septate, and more usually coloured, than in _Elvellacei_. Only a few solitary instances occur of individual species that are parasitic on living plants. [Illustration: FIG. 42.--_Sphaeria aquila._] In the _Sphaeriacei_, the substance of the stroma (when present) and of the perithecia is variable, being between fleshy and waxy in _Nectriei_, and tough, horny, sometimes brittle, in _Hypoxylon_. A perithecium, or cell excavated in the stroma which fulfils the functions of a perithecium, is always present. The hymenium lines the inner walls of the perithecium, and forms a gelatinous nucleus, consisting of asci and paraphyses. When fully mature, the asci are ruptured and the sporidia escape by a pore which occupies the apex of the perithecium. Sometimes the perithecia are solitary or scattered, and sometimes gregarious, whilst in other instances they are closely aggregated and immersed in a stroma of variable size and form. Conidia, spermatia, pycnidia, &c., have been traced to and associated with some species, but the history of others is still obscure. Many of the coniomycetous forms grouped under the _Sphaeronemei_ are probably conditions of the _Sphaeriacei_, as are also the _Melanconiei_, and some of the _Hy
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