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ch they have developed?" and answers the question with the astounding statement, "It seems reasonable to suppose that they have this power, it being simply a phase of heredity, the tendency of like to beget like." The right understanding of this famous problem is therefore fraught with the most important consequences to eugenics. The huge mass of experimental evidence that has been accumulated during the last quarter of a century has, necessarily, been almost wholly based on work with plants and lower animals. Even though we can not attempt to present a general review of this evidence, for which the reader must consult one of the standard works on biology or genetics, we shall point out some of the considerations underlying the problem and its solution. In the first place, it must be definitely understood that we are dealing only with specific, as distinguished from general, transmission. As the germ-cells derive their nourishment from the body, it is obvious that any cause profoundly affecting the latter might in that way exercise an influence on the germ-cells; that if the parent was starved, the germ-cells might be ill-nourished and the resulting offspring might be weak and puny. There is experimental evidence that this is the case; but that is not the inheritance of an acquired character. If, however, a white man tanned by long exposure to the tropical sun should have children who were brunettes, when the family stock was all blond; or if men whose legs were deformed through falls in childhood should have children whose legs, at birth, appeared deformed in the same manner; then there would be a distinct case of the transmission of an acquired characteristic. "The precise question," as Professor Thomson words it, "is this: Can a structural change in the body, induced by some change in use or disuse, or by a change in surrounding influence, affect the germ-cells in such a _specific_ or representative way that the offspring will through its inheritance exhibit, even in a slight degree, the modification which the parent acquired?" He then lists a number of current misunderstandings, which are so widespread that they deserve to be considered here. (1) It is frequently argued (as Herbert Spencer himself suggested) that unless modifications are inherited, there could be no such thing as evolution. Such pessimism is unwarranted. There _is_ abundant explanation of evolution, in the abundant supply of germinal variations
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