sive means of
modification.

1. VARIATION UNDER DOMESTICATION.

Causes of Variability. Effects of Habit. Correlation of Growth.
Inheritance. Character of Domestic Varieties. Difficulty of
distinguishing between Varieties and Species. Origin of Domestic
Varieties from one or more Species. Domestic Pigeons, their Differences
and Origin. Principle of Selection anciently followed, its Effects.
Methodical and Unconscious Selection. Unknown Origin of our Domestic
Productions. Circumstances favourable to Man's power of Selection.

When we look to the individuals of the same variety or sub-variety of
our older cultivated plants and animals, one of the first points which
strikes us, is, that they generally differ much more from each other,
than do the individuals of any one species or variety in a state of
nature. When we reflect on the vast diversity of the plants and animals
which have been cultivated, and which have varied during all ages under
the most different climates and treatment, I think we are driven to
conclude that this greater variability is simply due to our domestic
productions having been raised under conditions of life not so uniform
as, and somewhat different from, those to which the parent-species have
been exposed under nature. There is, also, I think, some probability
in the view propounded by Andrew Knight, that this variability may be
partly connected with excess of food. It seems pretty clear that organic
beings must be exposed during several generations to the new conditions
of life to cause any appreciable amount of variation; and that when the
organisation has once begun to vary, it generally continues to vary for
many generations. No case is on record of a variable being ceasing to be
variable under cultivation. Our oldest cultivated plants, such as wheat,
still often yield new varieties: our oldest domesticated animals are
still capable of rapid improvement or modification.

It has been disputed at what period of life the causes of variability,
whatever they may be, generally act; whether during the early or late
period of development of the embryo, or at the instant of conception.
Geoffroy St. Hilaire's experiments show that unnatural treatment of the
embryo causes monstrosities; and monstrosities cannot be separated by
any clear line of distinction from mere variations. But I am strongly
inclined to suspect that the most frequent cause of variability may
be attributed to the male and fem