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tally from mine in the belief that the primary germ-cell was formed within the ovarium of the female and was fertilised by the male. My gemmules are supposed to be formed, quite independently of sexual concourse, by each separate cell or unit throughout the body, and to be merely aggregated within the reproductive organs. Lastly, Mr. Herbert Spencer ('Principles of Biology,' vol. i., 1863-4, chaps. iv. and viii.) has discussed at considerable length what he designates as physiological units. These agree with my gemmules in being supposed to multiply and to be transmitted from parent to child; the sexual elements are supposed to serve merely as their vehicles; they are the efficient agents in all the forms of reproduction and in the repairs of injuries; they account for inheritance, but they are not brought to bear on reversion or atavism, and this is unintelligible to me; they are supposed to possess polarity, or, as I call it, affinity; and apparently they are believed to be derived from each separate part of the whole body. But gemmules differ from Mr. Spencer's physiological units, inasmuch as a certain number, or mass of them, are, as we shall see, requisite for the development of each cell or part. Nevertheless I should have concluded that Mr. Spencer's views were fundamentally the same with mine, had it not been for several passages which, as far as I understand them, indicate something quite different. I will quote some of these passages from pp. 254-256. "In the fertilised germ we have two groups of physiological units, slightly different in their structures."... "It is not obvious that change in the form of the part, caused by changed action, involves such change in the physiological units throughout the organism, that these, when groups of them are thrown off in the shape of reproductive centres, will unfold into organisms that have this part similarly changed in form. Indeed, when treating of Adaptation, we saw that an organ modified by increase or decrease of function can but slowly so react on the system at large as to bring about those correlative changes required to produce a new equilibrium; and yet only when such new equilibrium has been established, can we expect it to be _fully_ expressed in the modified physiological units of which the organism is built--only then can we count on a complete transfer of the modification to descendants."... "That the change in the offspring must, other things equal, be
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