r and with an accessory
fleshy bundle.
The hind limb of the Gorilla, therefore, ends in a true foot, with a
very moveable great toe. It is a prehensile foot, indeed, but is in no
sense a hand: it is a foot which differs from that of man not in
any fundamental character, but in mere proportions, in the degree of
mobility, and in the secondary arrangement of its parts.
It must not be supposed, however, because I speak of these differences
as not fundamental, that I wish to underrate their value. They are
important enough in their way, the structure of the foot being in strict
correlation with that of the rest of the organism in each case. Nor can
it be doubted that the greater division of physiological labour in Man,
so that the function of support is thrown wholly on the leg and foot, is
an advance in organization of very great moment to him; but, after all,
regarded anatomically, the resemblances between the foot of Man and
the foot of the Gorilla are far more striking and important than the
differences.
I have dwelt upon this point at length, because it is one regarding
which much delusion prevails; but I might have passed it over without
detriment to my argument, which only requires me to show that, be the
differences between the hand and foot of Man and those of the Gorilla
what they may--the differences between those of the Gorilla, and those
of the lower Apes are much greater.
It is not necessary to descend lower in the scale than the Orang for
conclusive evidence on this head.
The thumb of the Orang differs more from that of the Gorilla than
the thumb of the Gorilla differs from that of Man, not only by its
shortness, but by the absence of any special long flexor muscle. The
carpus of the Orang, like that of most lower apes, contains nine bones,
while in the Gorilla, as in Man and the Chimpanzee, there are only
eight.
The Orang's foot (Fig. 19) is still more aberrant; its very long
toes and short tarsus, short great toe, short and raised heel, great
obliquity of articulation in the leg, and absence of a long flexor
tendon to the great toe, separating it far more widely from the foot of
the Gorilla than the latter is separated from that of Man.
But, in some of the lower apes, the hand and foot diverge still more
from those of the Gorilla, than they do in the Orang. The thumb ceases
to be opposable in the American monkeys; is reduced to a mere rudiment
covered by the skin in the Spider Monkey; and is
|