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indrical sheaths. v, The dorsal "hood" formed by an enlargement in this region of the annular lobe of the fore-foot (m in figs. 2, 3). V, A swelling of the mantle-skirt, indicating the position on its inner face of the nidamental gland (see fig. 4, g.n.).] _Visceral Hump and Shell._--The visceral hump of _Nautilus_ (if we exclude from consideration the fine siphuncular pedicle which it trails, as it were, behind it) is very little, if at all, affected by the coiled form of the shell which it carries, since the animal always slips forward in the shell as it grows, and inhabits a chamber which is practically cylindrical (fig. 1). Were the deserted chambers thrown off instead of being accumulated behind the inhabited chamber as a coiled series of air-chambers, we should have a more correct indication in the shell of the extent and form of the animal's body. Amongst Gastropods it is not very unusual to find the animal slipping forward in its shell as growth advances and leaving an unoccupied chamber in the apex of the shell. This may indeed become shut off from the occupied cavity by a transverse septum, and a series of such septa may be formed, but in no Gastropod are these apical chambers known to contain a gas during the life of the animal in whose shell they occur. A further peculiarity of the nautilus shell and of that of the allied extinct _Ammonites_, _Scaphites_, _Orthoceras_, &c., and of the living _Spirula_, is that the series of deserted air-chambers is traversed by a cord-like pedicle extending from the centro-dorsal area of the visceral hump to the smallest and first-formed chamber of the series. No structure comparable to this siphuncular pedicle is known in any other Mollusca. The siphuncle does not communicate with the coelomic cavity; it is a simple vascular process of the mantle, whose cavity consists of a venous sinus, and whose wall contains a ramification of the pallial artery. There appears to be no doubt that the deserted chambers of the nautilus shell contain in the healthy living animal a gas which serves to lessen the specific gravity of the whole organism. This gas is said to be of the same composition as the atmosphere, with a larger proportion of nitrogen. With regard to its origin we have only conjectures. Each septum shutting off an air-containing chamber is formed during a period of quiescence, probably after the reproductive act, when the visceral mass of the nautilus may be sli
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